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On the first hot day of the year, midway through my six-mile run I turned back, into the wind, and both my heart rate and fatigue level skyrocketed, even as I was barely sweating—then, after the run, I sweated profusely. How can I stop that from happening again? – Charles
The only way to avoid bonking on the first hot day of the year is to slow down, hydrate, wear cool clothes, and pick a time of day when you won’t need shades. That said, a transition period of 7–14 days will not only acclimate you to the heat, it will leave you a stronger runner.
This happens every year—and in the age of climate change, sometimes multiple times per year. The weather shifts from cold to hot in the blink of an eye. Our bodies, not so much. The good news is a brief transition period of 7–14 days will prepare you for intense summer running. The bad news is your Strava posts are going to be a little underwhelming in the interim.
Understanding how heat affects your body begins with understanding how your body handles heat. When you run, your muscles produce energy. But your muscles aren’t very efficient at capturing this energy, using only 20%–25% and losing the rest as heat. The result is a rising core temperature. You can run effectively with a core temperature of about 100–102°F. Higher than that can become uncomfortable and even dangerous. So to avoid overheating, your muscles export lost heat energy by transferring it to your blood, which then carries it to your skin. Once at the skin, heat is offloaded through several ways, the two major ones being:
So far, so good. But what happens when the outside temperature soars? Well, on the first hot day of the year, you quickly reach a point where your body can’t offload heat quickly enough to offset your rising core temperature. During moderate effort runs, this effect kicks in at about 75°F. For more intense efforts, the outside temperature only needs to reach 65–70°F. At that point, this happens:
When all that occurs during a hard run on the first hot day of the year, you bonk. And don’t be misled if you seem to sweat less during the run than after you finish. That’s an illusion. During the run, air movement—a combination of your motion plus any wind—spurs faster evaporation. Once you stop running, the air movement stops (or slows), and ta-da!, you’re drenched.
So how do we avoid the misery described above? It begins with acknowledging you aren’t ready for the heat, then following these guidelines during a 1–2 week transition period:

And now for the best part. After a couple weeks of heat acclimatization, your body will undergo adaptations that make you a better runner! According to an Australian review, your blood plasma will increase by up to 12%, your heart rate will go down, your ventilation will go up, you’ll sweat more, and your energy requirements will be reduced.
With a few runs a week, your acclimatization should hold through the summer, during which you’ll be able to push yourself at will without ever re-experiencing that Day One bonk.